Henrik Kacser began studying for a diploma in animal genetics within the Institute of Animal Genetics in 1951, after gaining his doctorate in physical chemistry at Queen's University, Belfast. He made such a good impression that C.H. Waddington, at Beale's recommendation, asked Kacser if he would like to stay on as lecturer. He remained in Edinburgh for the rest of his life, working chiefly on physical and chemical approaches to biology, including metabolic control analysis.

This material was most likely gathered together as part of the research for Beale's obituary of Kacser, which he wrote with Grahame Bulfield for the Royal Society.